This research examines how food systems in the mountainous margins of northwest Nepal are being reshaped by climate change and increasing migration. Centering on the Dolpo region in Nepal’s northwestern Karnali province, the study explores how socioecological knowledge, practices, and relations involved in food provisioning are transformed within and between three interlinked agropastoral communities. Following rice, coarse grains, and local varietals through their production, processing, provisioning, and consumption, the project traces historical and contemporary social, economic, and environmental processes, including the afterlives of the grain-salt trade. It investigates how plains-grown rice shifted from a bartered delicacy to a daily commodity, and how efforts to marketize millet, buckwheat, and beans are refiguring values and practices in mountain agriculture. Ultimately, the study examines the relationship between rootedness, mobility, and value in agropastoral lifeways amid climatic uncertainty, migration, and the persistent devaluation of rural livelihoods.